"Rémy (Patton Oswalt) lives in a rat colony in the attic of a French country home with his brother Émile (Peter Sohn) and father Django (Brian Dennehy). Unlike his kin, Rémy is a gourmet whose keen sense of smell is used by the colony only to detect rat poison in food. But Rémy yearns for more, sneaking into the kitchen to read the cookbook of his hero, Parisian chef Auguste Gusteau (Brad Garrett), who appears to Rémy in visions throughout the film to expound on his motto that "anyone can cook." Rémy learns that Gusteau died after a harsh review from mean-spirited food critic Anton Ego (Peter O'Toole)."
The rats flee the house when the resident, an old woman, discovers the colony. Rémy, separated from the others, floats in the storm drains to Paris on Gusteau's cookbook, following the chef's image to his namesake restaurant, now run by former sous-chef Skinner (Ian Holm). As Rémy watches from a kitchen skylight, Alfredo Linguini (Lou Romano), a young man with no culinary talent, arrives and is hired on at the request of his recently deceased mother to do janitorial duties. The boy, unknown to all but his mother, Renata, is in fact Gusteau's son. Linguini spills a pot of soup and attempts to cover up his mistake by adding nearby random ingredients. Horrified by Linguini's actions, Rémy falls into the kitchen and though desperately trying to escape, cannot help but stop and attempt to fix the ruined soup. Rémy is caught in the act by Linguini, who himself is caught by Skinner as he captures the rat, but not before some of the soup has been served. To everyone's surprise, the soup is a success. The kitchen's sole woman cook, Colette (Janeane Garofalo), convinces Skinner not to fire Linguini provided he can recreate the soup. And thus begins an alliance, uneasy at first, by which Rémy secretly directs Linguini. The two perfect a marionette-like arrangement by which Rémy tugs at Linguini's hair to control his movements and stays hidden under Linguini's toque blanche."
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